Fossil evidence of biotic interactions is of considerable importance in understanding broad-scale evolutionary patterns and has long attracted the interest of paleontologists (Clarke, 1921; Moodie, 1923; for exhaustive compendium see Boucot, 1990). Although it is notoriously difficult to document with fossils, predation is ubiquitous in modern communities and often of fundamental importance in controlling the abundance and diversity of organisms in marine environments (e.g., Paine, 1969).